Main page


BENSCH, S., HELBIG, A.J., SALOMON, M., & SEIBOLD, I. 2002. Amplified fragment length polymorphism analysis identifies hybrids between two subspecies of warblers. Molecular Biology 11: 473-481.

BUTURLIN, S.A. & DEMENTIEV, G.P. 1937. [A complete key to the Birds of the USSR. vo. 4. Passeriformes.] Moscow-Leningrad. (In Russian.)

CLEMENT, P., HELBIG A.J & SMALL, B. 1998. Taxonomy and identification of chiffchaffs in the Western Palearctic. Brit. Birds 91: 361-376.

COLLINSON, J.M., SHANNON, T., ARCHER, P., ODIN, N., RIDDINGTON, R. & WALSH, P. 2013. Genetic analysis of migrant Siberian Chiffchaff in Britain and Ireland. Brit. Birds 106: 109-113.

CONWAY, G. 2005. Population composition of wintering Common Chiffchaffs in southern England. Brit. Birds 98: 427-428.

COPETE. J. L. & LÓPEZ, F. 2013. Identificatiön de subespecies en el mosquitero comün (Phylloscopus collybita collybita, P. c. abietinus, P. c. tristis, P. c. brevirostris, P. c. caucasicus y P. c. menzbieri). In: Rodríguez. N., Garcia, J. & Copete J. L. (eds). 2013. El mosquitero ibérico. Grupo Iberico de Anillamiento. Léon.

DEAN, A. 2013. How many Siberian Chifchaffs are reaching Britain? Brit. Birds 106: 282-283.

DEAN, A. 2018. Genetic studies of  'Siberian Chiffchaff'. Brit. Birds 111: 546 - 548.

—  , Bradshaw, C., Martin, J., Stoddart, A. & Walbridge, G. 2010. The status in Britain of 'Siberian Chiffchaff'. Brit. Birds 103: 320-338.

DEAN, A.R. 1985. Review of British status and identification of Greenish Warbler. Brit. Birds 78: 437-451.

 2007. Enigmatic ‘grey-and-white’ Common Chiffchaffs. Brit. Birds 100: 497-499.

— & Svensson, L. 2005. Siberian Chiffchaff revisited. Brit. Birds 98: 396-410.

de KNIJFF, P., van der SPEK, V. & FISHER, J. 2012. Genetic identity of grey chiffchaffs trapped in the Netherlands in autumns of 2009 - 2011. Dutch Birding 34: 386-392.

DUBOIS, P.J. & DUQUET, M. 2008. Further thoughts on Siberian Chiffchaffs. Brit. Birds 101: 149-150.

HARRISON, R. G. & LARSON, E. L. Hybridization, Introgression, and the Nature of Species Boundaries. J Hered 2014; 105: 795-809.

HELBIG, A.J., MARTENS, J., SEIBOLD, I., HENNING, F., SCHOTTLER, B. & WINK, M. 1996. Phylogeny and species limits in the Palearctic Chiffchaff Phylloscopus collybita complex: mitochondrial genetic differentiation and bioacoustic evidence. Ibis 138: 650-666.

IVANISTKII, V., ILINA, I. & MAROVA, I. 2020. The sounds between the strophes: different chiffchaff taxa perform different tret calls in their song. Bioacoustics DOI: 10.1080/09524622.2020.1796788.

JÄNNES, H. 2002. Calls of Eastern Vagrants (cd and booklet). Helsinki.

LINDHOLM, A. 2008. Mixed song of Chiffchaffs in Northern Russia. Alula 14: 108-115.

MAROVA, I.M. & ALEKSEEV, V.N. 2008. [The population structure and distribution of vocal dialects of the Chiffchaff (Phylloscopus collybita) in the southern Ural Mountains.] Publications of the South Ural State Reserve. Release 1. (In Russian)

—, FEDEROV,V.V., SHIPILINA, D.A., & ALEKSEEV, V. N. 2009. Genetic and vocal differentiation in hybrid zones of birds: Siberian and European Chiffchaffs (Phylloscopus [collybita] tristis and Ph. c. abietinus) in the Southern Urals. Doklady Biological Sciences 427: 384–386.

    & LEONOVICH, V.V. 1993. [Hybridization of Siberian and European Chiffchaffs in the zone of sympatry.] Proc. Zool. Mus. Moscow University 30: 147-164. (In Russian)

—, SHIPILINA, D. A., FEDEROV, V. V. & IVANITSKII, V. V. 2013.  Siberian and East European chiffchaffs: geographical distribution, morphological features, vocalization, phenomenon of mixed singing, and evidences of hybridization in sympatry zone. In: Rodríguez. N., Garcia, J. & Copete J. L. (eds). 2013. El mosquitero ibérico. Grupo Iberico de Anillamiento. Léon.

—, SHIPILINA, D., FEDEROV, V., ALEKSEEV, V. & IVANITSKII, V. 2017. Interaction between Common and Siberian Chiffchaff in a contact zone. Ornis Fennica 94: 66–81.

      —,  ILLYINA, I., KVARTALNOV, P., GRABOVSKY, V., BELOKON, M., SOLOVYOVA, E. & IVANITSKII, V. 2021. From the Bosporus to Kopet Dagh: Morphological, Genetic and BioacousticVariation in the Chiffchaff in Turkey, the Caucasus and Western Turkmenistan. Ardea 109: 1 - 16.

MARTENS, J. & MEINCKE, C. 1989. [The Siberian Chiffchaff Phylloscopus collybita tristis: Song and Reaction of a central European Population in field tests.] J. Orn. 130: 455–473. (In German)

MILLINGTON, R. 2006. Identification of a Siberian Chiffchaff in Norfolk. Birding World 19: 14-16.

MOTTEAU, V., CAMPBELL, O., SENFELD, T., SHANNON, T.J., COLLINSON, J.M. & LLOYD, S. 2022.  Occurrence of Common Chiffchaff Phylloscopus collybita subspecies menzbieri in the United Arab Emirates confirmed by genetic analysis. Brit. Birds 115: 500 - 509.

NOESKE, A. & DEAN, A.R. 2006. 'Siberian Chiffchaff' in Europe: photo forum. Publications / Articles,

RAKOVIĆ, M., NETO, J.M., LOPES, R.J., KOBLIK, E.A., FADEEV, I.V., LOHMAN, Y.V., AGHAYAN, S.A., BOANO, G., PAVIA, M., PERLMAN, Y., KIAT, Y., BEN DOV, A., COLLINSON, J.M., VOELKER, G. & DROVETSKI, S.V. 2019. Geographic patterns of mtDNA and Z-linked sequence variation in the Common Chiffchaff and the 'chiffchaff complex'. PLoS ONE 14(1): e0210268. (

ROSELAAR, C.S. & SHIRIHAI, H. in prep. Handbook of geographical variation and distribution of Palearctic birds.

SALOMON, M., BRIED, A.J., HELBIG, A.J., & RIOFRIO, J. 1997. Morphometric differentiation between male Common Chiffchaffs Phylloscopus [c.] collybita Vieillot, 1817, and Iberian Chiffchaffs P. [c.] brehmii Homeyer, 1871, in a secondary contact zone (Aves: Syllviidae). Zool. Anzeiger 236: 25–36.

SEVERTZOV, N.A. 1873. [Vertical and Horizontal Distribution of Turkestan Wildlife]. Turkestan. (In Russian)

SHIPILINA, D., SERBYN, M., IVANITSKII, V., MAROVA, I. &  BACKSTRÖM, N. 2017. Patterns of genetic, phenotypic, and acoustic variation across a chiffchaff (Phylloscopus collybita abietinus/tristis) hybrid zone. Ecology and Evolution 2017; 1–12.

SHIRIHAI, H. & SVENSSON, L. 2018. Handbook of Western Palearctic Birds: Passerines. vol. 1. London.

SNIGIREWSKI, S. 1931. [On range expansion of birds in the southern Ural-Mountains and new subspecies for this region.] J. Ornithol. 79: 57-66. (In German)

SUSHKIN, P.P. 1925. [List and distribution of birds of the Russian Altai and nearest parts of NW Mongolia, with a description of new or little-known forms.] Leningrad. (In Russian)

SVENSSON, L. 1992. Identification Guide to European Passerines. Fourth edn. Stockholm.

TALLA, V., KALSOOM, F., SHIPILINA, D., MAROVA, I. & BACKSTRÖM, N. 2017. Heterogeneous Patterns of Genetic Diversity and Differentiation in European and Siberian Chiffchaff (Phylloscopus collybita abietinus / P. tristis). G3 (Bethesda, Md.). 7. 10.1534/g3.117.300152.

TICEHURST, C. 1938. A Systematic Review of the Genus Phylloscopus (Willow-warblers or Leaf-warblers). London. 

VAN DEN BERG, A., & THE SOUND APPROACH. 2009. Calls, identification and taxonomy of Siberian Chiffchaff: an analysis. Dutch Birding 31: 79–85.

VAN DER SPEK, V. & DE KNIJFF, P. 2021. Migrating Chiffchaff taxa in The Netherlands:  a ten-year genetic study. Dutch Birding 43: 120 - 131.

VAURIE. C. 1954. Systematic notes on palearctic birds. No. 9. Sylviinae: the genus Phylloscopus. Amer. Nov. Mus. no. 1685

WILLIAMSON, K.1962. Identification for Ringers. No. 2. The Genus Phylloscopus. British Trust for Ornithology.

WU, C-I. 2001. The genic view of the process of speciation. J. Evol. Biol. 14: 851 - 865.